The vertex of a quadratic equation or parabola is the highest or lowest point of that equation. It lies on the plane of symmetry of the entire parabola as well; whatever lies on the left of the parabola is a complete mirror image of whatever is on the right. If you want to find the vertex of a quadratic equation, you can either use the vertex formula, or complete the square.
P.S. I found this off a website since I haven't done that yet... sorry!